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Ref-1 is a Trx-dependent endonuclease that facilitates AP-1 DNA-binding activity [88], while AP-1 is a heterodimer composed of proteins belonging to the c-Fos, c-Jun, ATF, and Maf families and manifests transcriptional activity through the regulation of gene expression in response to a variety of stimuli, including cytokines, growth factors, and oxidative stress [89]